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The common hippopotamus is semiaquatic, inhabiting rivers, lakes and mangrove swamps, where territorial bulls preside over a stretch of river and groups of five to thirty females and young. A rough evolutionary lineage can be traced from Eocene and Oligocene species: Anthracotherium and Elomeryx to the Miocene species Merycopotamus and Libycosaurus and the very latest anthracotheres in the Pliocene.